x1 = RanGDP_nuc x2 = RCC1_RanGDP x3 = GDP x4 = RCC1 x5 = RCC1_RanGTP x6 = RCC1_Ran x7 = GTP x8 = RanGTP_nuc x9 = RanGAP x10 = RanBP1 x11 = RanGTP_cy x12 = RanGTP_RanBP1 x13 = RanGDP_cy