k1 = r
k2 = K1
k3 = g1
k4 = beta1
k5 = d_u
k6 = d_m
k7 = delta_i1
k8 = d_i
k9 = a1_v
k10 = a1_i
k11 = a1_u
k12 = p_m1
k13 = r0_m1
k14 = r0_m2
k15 = r_u_m1
k16 = r_v_m2
k17 = d_e1
k18 = d_e2
k19 = p_m2
k20 = K2
k21 = g2
k22 = beta2
k23 = delta_i2
k24 = b
k25 = c
k26 = omega
k27 = delta_v
k28 = h_u
k29 = h_v
k30 = h_m
k31 = a2_u
k32 = Whole_organism