k1 = r
k2 = beta_1
k3 = eta
k4 = gamma
k5 = alpha_1
k6 = alpha_2
k7 = beta_2
k8 = alpha_3
k9 = sigma
k10 = delta
k11 = mu
k12 = rho
k13 = m
k14 = mu_1
k15 = rho_1
k16 = m_1
k17 = Whole_organism