k1 = a_1 k2 = a_2 k3 = a_3 k4 = b_1__1 k5 = b_2__1 k6 = b_3__1 k7 = e k8 = alpha_1 k9 = alpha_2 k10 = beta_1 k11 = beta_2 k12 = beta_3 k13 = u_1 k14 = u_2 k15 = compartment