k1 = ksE1 k2 = JM k3 = JE k4 = ksE2 k5 = kP1 k6 = JCD k7 = kP2 k8 = JCE k9 = kdE k10 = kRE k11 = ksmiR k12 = kdmiR k13 = ksM k14 = JS k15 = kdM1 k16 = kdM2 k17 = ksCD1 k18 = ksCD2 k19 = kdCD1 k20 = kdCD2 k21 = ksCE k22 = kdCE1 k23 = kdCE2 k24 = ksR k25 = kP k26 = JP k27 = kdR k28 = kdP k29 = kdRE k30 = compartment k31 = S