k1 = alpha1 k2 = alpha2 k3 = beta1 k4 = beta2 k5 = delta1 k6 = gamma2 k7 = s k8 = rho k9 = omega k10 = mu k11 = alpha3 k12 = gamma3 k13 = sigma1 k14 = sigma2 k15 = sigma3 k16 = v k17 = theta k18 = pi k19 = Tissue k20 = E