k1 = sum_R_smad_cyt
k2 = sum_R_smad_nuc
k3 = sum_smad4_cyt
k4 = sum_smad4_nuc
k5 = cytoplasm
k6 = nucleus
k7 = __lp_r2_KCAT
k8 = __lp_r2_K1
k9 = __lp_r3_k5nc
k10 = __lp_r3_k5cn
k11 = __lp_r4_k4nc
k12 = __lp_r4_k4cn
k13 = __lp_r5_k2a
k14 = __lp_r5_k2d
k15 = __lp_r6_k3
k16 = __lp_r7_k6d
k17 = __lp_r7_k6a
k18 = __lp_r8_Vmax7
k19 = __lp_r8_K7
k20 = __cm_k20
k21 = __cm_k21
k22 = __cm_k22
k23 = __cm_k23